Research Fellow in Applied Mathematics, Harvard University, U.S.A. 1969-70 Lecturer in Biology, Harvard University, U.S.A. 1970-71 Scientific Officer, Maharashtra Association for Cultivation of Science, Pune, India 1971-73 Assistant Professor, Centre for Theoretical Studies, Indian Institute of Science, Bangalore, India 1973-75 Associate Professor, Centre for Theoretical Studies, Indian Institute of Science, Bangalore, India 1975-81 Professor, Centre for Theoretical Studies, Indian Institute of Science, Bangalore, India 1981- Chairman, Centre for Ecological Sciences, Indian Institute of Science, Bangalore, India 1982-1992 Visiting Professor of Human Biology, Stanford University, Stanford, California, U.S.A. Spring 1991 Honorary Professor, Jawaharlal Nehru Centre for Advanced Scientific Research, Bangalore, India 1990- ASTRA Professor in Biological Sciences, Indian Institute of Science, Bangalore, India 1991-94 Dinstinguished Indo-American Lecturer, University of California, Berkeley, U.S.A. 1995